@import url('https://fonts.googleapis.com/css2?family=DM+Serif+Display:ital@0;1&family=Outfit:wght@300;400;500;600&family=DM+Mono:wght@400;500&display=swap');

:root {
  --navy:     #0A1628;
  --ocean:    #1B4F8A;
  --sapphire: #2E7BC4;
  --sky:      #5BA8E8;
  --ice:      #B8D8F5;
  --pale:     #E8F2FB;
  --gold:     #C8A94A;
  --gold-lt:  #F0D98A;
  --slate:    #3A4A5C;
  --off-white:#F8FAFC;
  --white:    #FFFFFF;
  --green:    #3A9A5C;
  --font-display: 'DM Serif Display', Georgia, serif;
  --font-body:    'Outfit', system-ui, sans-serif;
  --font-mono:    'DM Mono', 'Courier New', monospace;
}
*, *::before, *::after { box-sizing: border-box; margin: 0; padding: 0; }
html { scroll-behavior: smooth; }
body { font-family: var(--font-body); font-size: 17px; line-height: 1.75; color: var(--slate); background: var(--off-white); }

.species-hero { background: var(--navy); color: var(--white); padding: calc(76px + 3rem) 2rem 3.5rem; position: relative; overflow: hidden; }
.species-hero::before { content: ''; position: absolute; inset: 0; background: radial-gradient(ellipse at 70% 50%, rgba(46,123,196,0.2) 0%, transparent 65%); pointer-events: none; }
.hero-inner { max-width: 860px; margin: 0 auto; position: relative; }
.breadcrumb { font-family: var(--font-mono); font-size: 0.7rem; letter-spacing: 0.1em; text-transform: uppercase; color: var(--gold); margin-bottom: 1rem; }
.breadcrumb a { color: var(--gold); text-decoration: none; }
.breadcrumb a:hover { text-decoration: underline; }
.taxon-stamp { font-family: var(--font-mono); font-size: 0.75rem; letter-spacing: 0.08em; color: var(--sky); margin-bottom: 0.4rem; text-transform: uppercase; }
.species-hero h1 { font-family: var(--font-display); font-size: clamp(2rem, 4.5vw, 3.2rem); font-weight: 400; line-height: 1.15; margin-bottom: 0.4rem; }
.species-hero h1 em { font-style: italic; color: var(--ice); }
.common-name { color: var(--ice); font-size: 1rem; margin-bottom: 1.5rem; }
.hero-deck { font-size: 1.08rem; color: rgba(255,255,255,0.82); max-width: 680px; line-height: 1.6; margin-bottom: 2rem; }
.tag-row { display: flex; gap: 0.5rem; flex-wrap: wrap; }
.tag { font-family: var(--font-mono); font-size: 0.67rem; letter-spacing: 0.07em; text-transform: uppercase; padding: 0.28rem 0.72rem; border-radius: 20px; border: 1px solid rgba(91,168,232,0.4); color: var(--sky); }
.tag.pharma  { border-color: rgba(200,169,74,0.5); color: var(--gold); }
.tag.agritech{ border-color: rgba(58,154,92,0.5); color: #5ec47a; }

.hero-img-wrap { width: 100%; max-height: 480px; overflow: hidden; position: relative; background: var(--ocean); }
.hero-img-wrap img { width: 100%; height: 480px; object-fit: cover; display: block; opacity: 0.88; }
.hero-img-wrap .img-credit { position: absolute; bottom: 0.6rem; right: 0.8rem; font-family: var(--font-mono); font-size: 0.62rem; color: rgba(255,255,255,0.55); }

.stat-bar { background: var(--ocean); border-bottom: 2px solid var(--gold); display: grid; grid-template-columns: repeat(auto-fit, minmax(140px,1fr)); }
.stat-item { padding: 1.2rem 1.5rem; text-align: center; border-right: 1px solid rgba(255,255,255,0.08); }
.stat-item:last-child { border-right: none; }
.stat-num { font-family: var(--font-display); font-size: 1.8rem; color: var(--gold-lt); display: block; line-height: 1; }
.stat-label { font-size: 0.72rem; color: var(--ice); letter-spacing: 0.04em; margin-top: 0.25rem; display: block; }

.article-wrap { max-width: 820px; margin: 0 auto; padding: 3rem 2rem 5rem; }
.article-wrap h2 { font-family: var(--font-display); font-size: 1.6rem; color: var(--navy); font-weight: 400; margin: 2.8rem 0 0.9rem; line-height: 1.25; }
.article-wrap h3 { font-size: 0.82rem; font-weight: 600; color: var(--ocean); text-transform: uppercase; letter-spacing: 0.07em; margin: 2rem 0 0.6rem; font-family: var(--font-mono); }
.article-wrap p { margin-bottom: 1.1rem; }
.article-wrap em { font-style: italic; }
.article-wrap strong { font-weight: 600; color: var(--navy); }

.callout { background: var(--pale); border-left: 4px solid var(--ocean); padding: 1.2rem 1.5rem; margin: 2rem 0; border-radius: 0 6px 6px 0; }
.callout p { margin: 0; color: var(--navy); font-size: 1.02rem; line-height: 1.6; }

.trait-table { width: 100%; border-collapse: collapse; margin: 2rem 0; font-size: 0.9rem; }
.trait-table th { background: var(--ocean); color: var(--white); padding: 0.65rem 1rem; text-align: left; font-family: var(--font-mono); font-size: 0.72rem; letter-spacing: 0.05em; text-transform: uppercase; font-weight: 500; }
.trait-table td { padding: 0.6rem 1rem; border-bottom: 1px solid var(--ice); }
.trait-table tr:nth-child(even) td { background: var(--pale); }

.crosslink-box { background: var(--navy); padding: 2rem; border-radius: 8px; margin: 3rem 0; }
.crosslink-box .cl-label { font-family: var(--font-mono); font-size: 0.72rem; letter-spacing: 0.09em; text-transform: uppercase; color: var(--gold); margin-bottom: 0.6rem; display: block; }
.crosslink-box p { color: var(--ice); font-size: 0.95rem; margin-bottom: 1.2rem; }
.cl-grid { display: grid; grid-template-columns: repeat(auto-fit, minmax(190px,1fr)); gap: 0.7rem; }
.cl-grid a { display: block; padding: 0.7rem 1rem; background: rgba(91,168,232,0.08); border: 1px solid rgba(91,168,232,0.22); border-radius: 6px; color: var(--sky); text-decoration: none; font-size: 0.88rem; transition: background 0.15s; }
.cl-grid a:hover { background: rgba(91,168,232,0.18); }
.cl-grid a .cl-type { font-family: var(--font-mono); font-size: 0.65rem; color: var(--gold); display: block; margin-bottom: 0.15rem; letter-spacing: 0.06em; text-transform: uppercase; }

.schema-stamp { margin-top: 3rem; padding-top: 1.5rem; border-top: 1px solid var(--ice); font-family: var(--font-mono); font-size: 0.7rem; color: #8a9bb0; display: flex; gap: 2rem; flex-wrap: wrap; }

/* INDEX */
.index-hero { background: var(--navy); padding: 4.5rem 2rem 3rem; text-align: center; }
.index-hero .eyebrow { font-family: var(--font-mono); font-size: 0.72rem; letter-spacing: 0.12em; text-transform: uppercase; color: var(--gold); margin-bottom: 0.75rem; }
.index-hero h1 { font-family: var(--font-display); font-size: clamp(2rem, 4vw, 3rem); color: var(--white); font-weight: 400; margin-bottom: 1rem; line-height: 1.2; }
.index-hero p { color: var(--ice); max-width: 600px; margin: 0 auto; font-size: 1.05rem; }

.filter-bar { background: var(--ocean); padding: 0.9rem 2rem; display: flex; gap: 0.5rem; flex-wrap: wrap; justify-content: center; border-bottom: 2px solid var(--gold); }
.filter-btn { font-family: var(--font-mono); font-size: 0.7rem; letter-spacing: 0.07em; text-transform: uppercase; padding: 0.38rem 1rem; border: 1px solid rgba(255,255,255,0.3); background: transparent; color: var(--ice); border-radius: 20px; cursor: pointer; transition: all 0.15s; }
.filter-btn:hover, .filter-btn.active { background: rgba(91,168,232,0.2); border-color: var(--sky); color: var(--sky); }

.species-grid { display: grid; grid-template-columns: repeat(auto-fill, minmax(300px,1fr)); gap: 1.5rem; padding: 2.5rem 2rem 4rem; max-width: 1200px; margin: 0 auto; }
.species-card { background: var(--white); border-radius: 10px; overflow: hidden; box-shadow: 0 2px 12px rgba(10,22,40,0.08); transition: transform 0.2s, box-shadow 0.2s; text-decoration: none; display: flex; flex-direction: column; }
.species-card:hover { transform: translateY(-4px); box-shadow: 0 8px 28px rgba(10,22,40,0.14); }
.card-img-wrap { width: 100%; height: 200px; overflow: hidden; background: var(--ocean); position: relative; }
.card-img-wrap img { width: 100%; height: 100%; object-fit: cover; display: block; }
.card-body { padding: 1.3rem 1.4rem; flex: 1; display: flex; flex-direction: column; }
.card-ecoregion { font-family: var(--font-mono); font-size: 0.65rem; letter-spacing: 0.09em; text-transform: uppercase; color: var(--sapphire); margin-bottom: 0.3rem; }
.card-taxon { font-family: var(--font-display); font-size: 1.15rem; color: var(--navy); font-weight: 400; margin-bottom: 0.15rem; font-style: italic; }
.card-common { font-size: 0.85rem; color: var(--slate); margin-bottom: 0.8rem; }
.card-hook { font-size: 0.9rem; color: var(--slate); line-height: 1.55; flex: 1; }
.card-tags { display: flex; gap: 0.4rem; flex-wrap: wrap; margin-top: 1rem; }
.card-tag { font-family: var(--font-mono); font-size: 0.63rem; letter-spacing: 0.06em; text-transform: uppercase; padding: 0.22rem 0.6rem; border-radius: 20px; border: 1px solid rgba(91,168,232,0.35); color: var(--sapphire); }
.card-tag.pharma  { border-color: rgba(200,169,74,0.5); color: var(--gold); }
.card-tag.agritech{ border-color: rgba(58,154,92,0.5); color: #3A9A5C; }

/* NAV DROPDOWN */
.has-dropdown { position: relative; }
.dropdown-menu { display: none; position: absolute; top: 100%; left: 0; background: var(--navy); border: 1px solid rgba(91,168,232,0.2); border-radius: 6px; min-width: 220px; padding: 0.5rem 0; z-index: 200; box-shadow: 0 8px 24px rgba(0,0,0,0.3); }
.has-dropdown:hover .dropdown-menu { display: block; }
.dropdown-menu a { display: block; padding: 0.5rem 1rem; color: var(--ice); text-decoration: none; font-size: 0.82rem; transition: background 0.12s; }
.dropdown-menu a:hover { background: rgba(91,168,232,0.1); color: var(--sky); }
.dropdown-menu .dd-section { font-family: var(--font-mono); font-size: 0.62rem; letter-spacing: 0.1em; text-transform: uppercase; color: var(--gold); padding: 0.6rem 1rem 0.3rem; }